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Asplenium trichomanes subsp. densum

(Brack.) W.H.Wagner (1999)

This name is accepted

Kingdom: Viridiplantae Phylum: Pteridophyta Class/Clade: Polypodiopsida Order: Polypodiales Family: Aspleniaceae Genus: Asplenium

‘oāli‘i, ‘owāli‘i, maidenhair spleenwort

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Description

Key Characters:

Growth Form:

Stems:

Roots:

Leaves:

Sori:

Gametophyte:

Ploidy:

Habitat:

Elevation Range:

Historical Distribution

Uses and Culture

USES

Natural History

Statewide Status

Endemic

Island Status

Kaua'i Endemic
O'ahu Unknown
Maui Endemic
Hawai'i Endemic

Dispersal Agents


Bibliography

Name Published In: Contr. Univ. Michigan Herb. 22: 160 (1999)

Other References

Palmer 2003:74 (EM, H); Faccenda, Arthur, et al. 2025:158 (O? [observed but uncollected])

Occurrences

SNo. Scientific Name Locality Habitat Basis of Record Description Recorded By Record Number Island Source Date
1 Asplenium trichomanes subsp. densum 2 mi NE of Huehue Ranch House, Kaupulehu, N. Kona in cracks on bare aa lava Preserved_Specimen St.John, H. 22447 Hawaii BISH 1946-12-28
2 Asplenium trichomanes subsp. densum Puuwaawaa, on transect T19 bare aa; with Metrosideros Preserved_Specimen terrestrial Takeuchi, W.N. 5835 Hawaii BISH 1989-05-01
3 Asplenium trichomanes subsp. densum Mauna Kea, Wai Kahalulu Gulch adz quarry Preserved_Specimen Howarth, F.G. Hawaii BISH 1991-08-10
4 Asplenium trichomanes subsp. densum above Halealoha in cave in 1850 flow Preserved_Specimen Forbes, C.N. 782 Hawaii BISH 1915-06-09
5 Asplenium trichomanes subsp. densum East Maui, Haleakala near summit Preserved_Specimen Fern mixed with Luzula Forbes, C.N. 214 Maui BISH 1910-07-01
6 Asplenium trichomanes subsp. densum East Maui, Haleakala cleft of rocks Preserved_Specimen St.John, H. 10349 Maui BISH 1930-02-15
7 Asplenium trichomanes subsp. densum Mauna Kea Preserved_Specimen U.S. Exploring Expedition 15 Hawaii BISH 1840-01-01
8 Asplenium trichomanes subsp. densum N. Kona, Kaniku lava field, crevices of aa clinker; with Doryopteris decora, Peperomia leptostachya, Metrosideros polymorpha, Myrsine lanaiensis, Xylosma hawaiiense var. hillebrandii, Colubrina oppositifolia Preserved_Specimen Takeuchi, W.N. 3540 Hawaii BISH 1987-05-01
9 Asplenium trichomanes subsp. densum Ukulele Preserved_Specimen Forbes, C.N. 726 Maui BISH 1919-07-11
10 Asplenium trichomanes subsp. densum Hualalai on summit plateau Preserved_Specimen Neal, M.C. Hawaii BISH 1934-06-19
11 Asplenium trichomanes subsp. densum Hawaii Preserved_Specimen Rémy, J. 22 Hawaii BISH 1851-01-01
12 Asplenium trichomanes subsp. densum above Mauna Kea-Humuula trail Preserved_Specimen Hartt, C.E. 751 Hawaii BISH 1935-08-09
13 Asplenium trichomanes subsp. densum Maui Preserved_Specimen Brigham, W.T. Maui BISH 1909-01-01
14 Asplenium trichomanes subsp. densum Below Puu Oo Preserved_Specimen Forbes, C.N. 812 Hawaii BISH 1915-06-11
15 Asplenium trichomanes subsp. densum road to Haleakala summit Preserved_Specimen Palmer, M.R. 98 Maui BISH 1955-12-23
16 Asplenium trichomanes subsp. densum Below Puu Oo Preserved_Specimen Forbes, C.N. 812 Hawaii BISH 1915-06-11
17 Asplenium trichomanes subsp. densum Waikaloa Preserved_Specimen Hobdy, R.W. 2449 Hawaii BISH 1985-09-12
18 Asplenium trichomanes subsp. densum Puu Laau, near hunter's cabin Preserved_Specimen weed Herbst, D.R. 5204 Hawaii BISH 1975-01-18
19 Asplenium trichomanes subsp. densum Along Saddle Road, Pohakuloa Training Area, between 30 and 45 mile markers, within 60 feet of the road. Preserved_Specimen Funk, E.J. 57 Hawaii BISH 1990-12-13
20 Asplenium trichomanes subsp. densum Mauna Kea, above Humuula Preserved_Specimen Neal, M.C. 1010 Hawaii BISH 1935-08-15
21 Asplenium trichomanes subsp. densum East Maui, crater of Haleakala Preserved_Specimen Forbes, C.N. 306 Maui BISH 1910-08-01
22 Asplenium trichomanes subsp. densum East Maui, Haleakala, summit Preserved_Specimen Munro, G.C. 659 Maui BISH 1918-04-24
23 Asplenium trichomanes subsp. densum S slope of Haleakala, Puu Pane Preserved_Specimen Forbes, C.N. 1900 Maui BISH 1920-03-08
24 Asplenium trichomanes subsp. densum Mauna Loa; E. slopes at road from Kilauea 19.4N 155.3W Preserved_Specimen Krajina, V.J. 611226059 Hawaii BISH 1961-12-26
25 Asplenium trichomanes subsp. densum Below Puu Oo Preserved_Specimen Forbes, C.N. 812 Hawaii BISH 1915-06-11
26 Asplenium trichomanes subsp. densum Puuwaawaa Preserved_Specimen Forbes, C.N. 27 Hawaii BISH 1911-06-09
27 Asplenium trichomanes subsp. densum Mauna Kea, Waikahalulu Gulch Preserved_Specimen McEldowney, H. 23 Hawaii BISH 1976-09-01
28 Asplenium trichomanes subsp. densum East Maui, Kalapawili Grasslands Preserved_Specimen Welton, P. 1989 Maui BISH 1997-09-26
29 Asplenium trichomanes subsp. densum East Maui, Haleakala, near summit Preserved_Specimen Munro, G.C. 451 Maui BISH 1918-06-26
30 Asplenium trichomanes subsp. densum National Park, near observatory at end of Mauna Loa truck road Preserved_Specimen Webster, G.L. 1733 Hawaii BISH 1948-06-16
31 Asplenium trichomanes subsp. densum Between Glenwood and Volcano House Preserved_Specimen Degener, O. 30439 Hawaii BISH 1926-07-27
32 Asplenium trichomanes subsp. densum Kona Preserved_Specimen Smith, F.G. 49 Hawaii BISH 1929-03-15
33 Asplenium trichomanes subsp. densum Mauna Kea, Wai Kahalulu Gulch Preserved_Specimen Howarth, F.G. Hawaii BISH 1991-08-10
34 Asplenium trichomanes subsp. densum Hualalai Preserved_Specimen Meebold, A. Hawaii BISH 1932-05-01
35 Asplenium trichomanes subsp. densum Maui Preserved_Specimen (Not on sheet) Maui BISH
36 Asplenium trichomanes subsp. densum East Maui, Haleakala Preserved_Specimen Anderson, D. Maui BISH 1932-08-20
37 Asplenium trichomanes subsp. densum Keawewai, Keauhou Preserved_Specimen Dillingham, M. Hawaii BISH 1980-12-04
38 Asplenium trichomanes subsp. densum Maui Preserved_Specimen (Not on sheet) Maui BISH
39 Asplenium trichomanes subsp. densum East Maui, Haleakala Preserved_Specimen Anderson, D. Maui BISH 1932-08-20
40 Asplenium trichomanes subsp. densum Volcano Kilauea Preserved_Specimen Forbes, C.N. Hawaii BISH 1908-09-01
41 Asplenium trichomanes subsp. densum Saddle Rd Preserved_Specimen Woolford, H.A. Hawaii BISH 1954-12-20
42 Asplenium trichomanes subsp. densum Kilauea, Bird Park Preserved_Specimen Neal, M.C. Hawaii BISH 1929-08-05
43 Asplenium trichomanes subsp. densum Kona; Keauhou No.1; saddle road between Hualalai and Mauna Loa Preserved_Specimen Christ, J.H. Hawaii BISH 1958-05-15
44 Asplenium trichomanes subsp. densum Oahu or Hawaii (Kona or Kilauea) Preserved_Specimen Lack, F. BISH 1880-01-01
45 Asplenium trichomanes subsp. densum unknown Preserved_Specimen Baldwin, D.D. 113 BISH 1875-01-01
46 Asplenium trichomanes subsp. densum unknown Preserved_Specimen Forbes, C.N. BISH 1908-11-14
47 Asplenium trichomanes subsp. densum Haleakala, 2 mi W of Paliku cabins crater floor in aa lava, with Pellaea ternifolia Preserved_Specimen Henrickson, J.S. 3713 Maui BISH 1969-06-24
48 Asplenium trichomanes subsp. densum unknown Preserved_Specimen (Not on sheet) BISH
49 Asplenium trichomanes subsp. densum unknown Preserved_Specimen Hillebrand, W.B. BISH
50 Asplenium trichomanes subsp. densum Puu Hualalai, Puuwaawaa crevices of lava Preserved_Specimen St.John, H. 11412 Hawaii BISH 1931-12-30
51 Asplenium trichomanes subsp. densum Hualalai, location 17 edge of koa forest Preserved_Specimen Kondo, Y. Hawaii BISH 1946-07-10
52 Asplenium trichomanes subsp. densum W outer edge of Haleakala, below Kalahaku Overlook alpine scrubland on rock slopes; with Sophora chrysophylla, Geranium tridens, Styphelia tameiameiae, Coprosma montana, Vaccinium, Dodonaea eriocarpa, Silene struthioloides, Tetramalopium humile Preserved_Specimen Iltis, H.H. 555 Maui BISH 1967-08-07
53 Asplenium trichomanes subsp. densum Makawao District, Haleakala National Park At base of north-facing cliff face, 1/2 mile south of Holua cabin. Preserved_Specimen Sun and shade fronds. Bruegmann, M.M. 184 Maui BISH 1987-01-30
54 Asplenium trichomanes subsp. densum Hualalai in cave; on summit plateau Preserved_Specimen Neal, M.C. Hawaii BISH 1934-06-19
55 Asplenium trichomanes subsp. densum Saddle Road, Mauna Kea side, 23.5 miles from Hilo More than 10 plants growing in dim light within 20 ft of lava tube entrance, on rocks with mosses and liverworts. Open 'ohi'a forest with Styphelia, Vaccinium spp, Coprosma ernodeoides, Asplenium adiantum-nigrum & Machaerina spp. Preserved_Specimen Davis, J. 529 Hawaii BISH 1981-06-15
56 Asplenium trichomanes subsp. densum National Park, beginning of Mauna Loa strip road, off the trail to weather station lava rock, open sunny with grasses Preserved_Specimen Herat, T. 618 Hawaii BISH 1974-06-19
57 Asplenium trichomanes subsp. densum Rest House, rim of Haleakala Crater under overhanging rocks Preserved_Specimen Fosberg, F.R. 9960 Maui BISH 1933-08-21
58 Asplenium trichomanes subsp. densum Hawaii Table land. Preserved_Specimen Mann, H. 257 Hawaii BISH 1864-01-01
59 Asplenium trichomanes subsp. densum Kona; Mauka of Papaloa; Headquarters of Kealakekua Ranch With Scrubby Ohia and Mamane Preserved_Specimen Christ, J.H. Hawaii BISH 1958-11-23
60 Asplenium trichomanes subsp. densum Hamakua, Waimea, Upper Paauhau rock crevices Preserved_Specimen Hosaka, E.Y. 2199 Hawaii BISH 1938-06-03
61 Asplenium trichomanes subsp. densum Kiilae, S. Kona kipuka in pahoehoe crevice Preserved_Specimen Hosaka, E.Y. 1522 Hawaii BISH 1936-08-21
62 Asplenium trichomanes subsp. densum 6 mi up narrow paved road toward Mauna Loa from jct. with rte 20; turnoff just E of Puu Huluhulu tan pahoehoe lava Preserved_Specimen Stuessy, T.F. 6156 Hawaii BISH 1982-05-30
63 Asplenium trichomanes subsp. densum ~23 mi post, Saddle Road crevices in barren aa lava Preserved_Specimen Takeuchi, W.N. 1 Hawaii BISH 1983-11-26
64 Asplenium trichomanes subsp. densum Mauna Kea, Wai Kahalulu Adz Quarry, shelter cave Preserved_Specimen Stone, F.D. Hawaii BISH 1991-08-10
65 Asplenium trichomanes subsp. densum East Maui, Hana District, Crater of Haleakala, Paliku open stand of Sophora and Myrsine under mostly tree branch Preserved_Specimen St.John, H. 20974 Maui BISH 1945-08-12
66 Asplenium trichomanes subsp. densum Mauna Loa Strip Rd, Keamoku Flow Recent lava flow (early 1980's); cracks in rough bare lava surface. Preserved_Specimen Tufted fern with blackish stipes and erect fronds. Sykes, W.R. 31591 Hawaii BISH 1991-06-02
67 Asplenium trichomanes subsp. densum Olinda, Kula pipe line by stream, on rocks Preserved_Specimen St.John, H. 10297 Maui BISH 1930-02-11
68 Asplenium trichomanes subsp. densum East Maui, N outer slope of Haleakala, 1.5 mi N of Paliku Cabins, NW of Puu Alaea small shaded gulch, lower heath scrub, with Sadleria, Vaccinium, Dryopteris Preserved_Specimen Henrickson, J.S. 3677 Maui BISH 1969-06-22
69 Asplenium trichomanes subsp. densum Kona; Kealakekua Ranch; Near Puu O Uo In `a`a lava; Scattered Colonies of Vegetation. Preserved_Specimen Christ, J.H. Hawaii BISH 1958-11-23
70 Asplenium trichomanes subsp. densum Hawaii Table land. Preserved_Specimen Mann, H. 257 Hawaii BISH 1864-01-01
71 Asplenium trichomanes subsp. densum Puu Huluhulu, Humuula, N. Hilo open woods on hill kipuka, in crevice of tree bark Preserved_Specimen St.John, H. 23941 Hawaii BISH 1949-12-24
72 Asplenium trichomanes subsp. densum W slope of Haleakala with Santalum haleakalae Preserved_Specimen Gillett, G.W. 1560 Maui BISH 1964-04-17
73 Asplenium trichomanes subsp. densum Pohakuloa Military Camp, Mauna Kea 1979 burn area. Preserved_Specimen Adee, K.T. Hawaii BISH
74 Asplenium trichomanes subsp. densum 6.75 mi E of Puu Huluhulu, Saddle Rd mossy forest of Metrosideros in kipuka Preserved_Specimen St.John, H. 25346 Hawaii BISH 1954-12-20
75 Asplenium trichomanes subsp. densum Haleakala, 2 mi W of Paliku cabins crater floor in aa lava, with Pellaea ternifolia Preserved_Specimen Henrickson, J.S. 3713 Maui BISH 1969-06-24
76 Asplenium trichomanes subsp. densum East Maui, Haleakala Crater, Halemauu Trail rock crevices Preserved_Specimen Fosberg, F.R. 9987 Maui BISH 1933-08-23
77 Asplenium trichomanes subsp. densum East Maui, Koolau Gap, Waikau base of pali, under shaded ledge Preserved_Specimen large Hobdy, R.W. 2285 Maui BISH 1985-02-09
78 Asplenium trichomanes subsp. densum SW side of Haleakala, Puu Ouli and above on aa lava Preserved_Specimen in thick clumps Forbes, C.N. 2149 Maui BISH 1920-04-05
79 Asplenium trichomanes subsp. densum SW flank of Mauna Loa, Manuka open Metrosideros on cinder, Machaerina, Coprosma, Styphelia, Vaccinium Preserved_Specimen Higashino, P.K. 6208 Hawaii BISH 1977-07-21
80 Asplenium trichomanes subsp. densum Hualalai lava fields near summit Preserved_Specimen Rock, J.F.C. 3615 Hawaii BISH 1909-06-09
81 Asplenium trichomanes subsp. densum National Park, Kipuka Ki on aa lava Preserved_Specimen Webster, G.L. 1717 Hawaii BISH 1948-06-16
82 Asplenium trichomanes subsp. densum East Maui, Hana District, Crater of Haleakala, foot of pali E of Waikekeehia in deep roch crevice on lava flow Preserved_Specimen St.John, H. 17773 Maui BISH 1936-12-25
83 Asplenium trichomanes subsp. densum Humuula, base of Puu Huluhulu on lava flow at base Preserved_Specimen Neal, M.C. 664 Hawaii BISH 1935-08-06
84 Asplenium trichomanes subsp. densum Kau, Manuka moist ohia forest, shaded Preserved_Specimen 10 terrestrial Russ, G.W. 47 Hawaii BISH 1927-07-11
85 Asplenium trichomanes subsp. densum East Maui, Koolau Forest Reserve in lava crevices in gulch in tussock grasslands with other ferns, lichens and mosses Preserved_Specimen Harrison, B. 508 Maui BISH 1973-11-09
86 Asplenium trichomanes subsp. densum Along Road 8 1/2 miles North of Volcano House; Kilauea, Hawaii Kipuka Preserved_Specimen Degener, O. 30438 Hawaii BISH 1926-07-22
87 Asplenium trichomanes subsp. densum 18.6 mi W of Hilo, Rte 20 aa on both sides of road Preserved_Specimen clumps up to 15 cm with many fronds Stuessy, T.F. 6133 Hawaii BISH 1982-05-30
88 Asplenium trichomanes subsp. densum East Maui, W fork of W Wailua Iki Montane shrubland dominated by Sadleria cyatheoides. W/ scattered Metrosideros polymorpha incana and shrubs of Leptecophylla, Coprosma ernodeoides, C. montana, Rubus hawaiensis, Dubautia reticulata, Morelotia gahniiformis, w/ riparian vegetation of bryophytes . Preserved_Specimen Wood, K.R. 13390 Maui BISH 2008-10-22
89 Asplenium trichomanes subsp. densum 0.4 mi SSE of Mauna Kea State Park, W side of Saddle Road open Myoporum, Sophora woodland; with exotic herbaceous species, Chenopodium oahuense, Dodonaea Preserved_Specimen Wagner, W.L. 4702 Hawaii BISH 1982-11-18
90 Asplenium trichomanes subsp. densum 3/4 miles down sliding sand trails( from obervation house) in crevice under lava rock ledge Preserved_Specimen Wilson, K.A. 63 Maui BISH 1952-12-24
91 Asplenium trichomanes subsp. densum 6.75 mi E of Puu Huluhulu, Saddle Rd kipuka Preserved_Specimen St.John, H. 25346 Hawaii BISH 1954-12-20
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